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![PDF] Tinbergen's four questions, organized: a response to Bateson and Laland. | Semantic Scholar](https://d3i71xaburhd42.cloudfront.net/6117ba207f35dbb9ef1c3c28d478e6b281e13d07/1-Table1-1.png "PDF] Tinbergen's four questions, organized: a response to Bateson and Laland. | Semantic Scholar")